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The mechanisms behind this variation are largely unclear, but may be partly related to Sex hamel in Norway drivers and density dependence operating differently on the adult male and female segments of the population. There were a positive population size trend and an overall female bias in ASR throughout the study period. Increased winter precipitation, a proxy for the harshness of winter feeding conditions, was found to significantly reduce adult population Sex hamel in Norway rates through reduced survival in males, but not in females.
However, increased population size tended to cause a stronger immediate decline in female than in male adult population growth rates. As a consequence, the female bias in ASR increased with harsher winter conditions and declined with higher population Flirting Trondheim. As expected from the increased frequency of rainy and icy winters due to climate warming, a recent trend toward increased female bias in ASR was evident.
In contrast, ASR in wild populations is frequently skewed and, in ungulates, generally toward females Berger and Gompper The observed skew in ASR results mainly from a higher average mortality in the rarer sex.
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The wild Svalbard reindeer Rangifer tarandus platyrhynchus is hamdl ideal model species to study the effects of climate and population size on demography. The endemic Svalbard reindeer, along with Svalbard rock ptarmigan Lagopus muta hyperboreaare the only overwintering native herbivores in Svalbard.
In addition, there is Sex hamel in Norway local population of the introduced sibling vole Microtus leviswhich only occurs in a small area close to a bird colony. The effect of interspecific resource competition with migratory geese on reindeer population dynamics is considered to be low due to the seasonal and local occurrence of geese. Reindeer harvesting in the study area is restricted to only parts of RD and Colesdalen. The regulations related to the hunting of Svalbard reindeer state that the harvest should not affect the population development or the population demography.
In line with this goal, sex and age distribution of annual harvest quotas aim to reflect the demographic structure of the population. Annual population censuses of Svalbard reindeer were Cabanas en Drobak Norway in summer after calving. Nirway two latter age classes were also categorized by sex.